>so when we are discussing Natural
Selection I have in view colonies that are truly unmanaged (or as they now
style them across the pond - 'free living').

I have not come across the term 'free living' before so not sure where that
comes from, but I would suggest that almost all feral colonies are not
really truly unmanaged as they are swarms that have escaped from managed
colonies and have therefore been selected before they swarmed.  A truly
feral, unmanaged colony must be a very rare thing these days, certainly in
most of the UK.

Best wishes

Peter 
52°14'44.44"N, 1°50'35"W

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