> The thing that is most amazing to me is that it's taking this long for researchers to learn the basic biology of this economically-important parasite.

Perhaps this recent overview is useful in this context (especially the references cited).

Biology and Management of Varroa destructor Mesostigmata: Varroidae) in Apis mellifera
(Hymenoptera: Apidae) Colonies
Morgan A. Roth, 1;  James M. Wilson,  2; Keith R. Tignor, 3 and Aaron D. Gross 1,4
1 Molecular Physiology and Toxicology Laboratory, Department of Entomology, Virginia Polytechnic Institute and State University, 307 Latham Hall (MC0390), 220 Ag Quad Ln., Blacksburg, VA 24061, 
2 Department of Entomology, Virginia Polytechnic and State University, Blacksburg, VA 24061, 
3 Office of Plant Industry Services, VA Department of Agriculture and Consumer Services, Richmond, VA, and 
4 Corresponding author, e-mail: [log in to unmask] Subject Editor: Matthew Messenger
Received 20 September 2019; Editorial decision 29 November 2019.

https://academic.oup.com/jipm/article/11/1/pmz036/5692075

Open access.

Kind regards,

Ghislain De Roeck,
Belgium.

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