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The modern experimental period of study on contact chemoreceptors of insects dates from the work of Minnich (1921, 1922), in which he demonstrated the presence of tarsal contact chemoreceptors in the butterflies, Vanessa atalanta and Aglais antiopa. ... The tarsal receptors of butterflies, he found, enable the insects to distinguish sugar solutions and other solutions from water. They, therefore, seem to be common chemical receptors at least, and probably truly gustatory. Here, at last, the search for the taste-organs of insects was broadened and put on a firm experimental basis -- Hubert Frings and Mable Frings (1949) Pennsylvania State College, State College, Pennsylvania
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