Capped brood is not a reliable indicator of loss.  Whether  environmental 
(e.g. cold chill), queen quality, nutrition, pesticide, viral) -  the queens 
often make up for losses of early developmental stages by quickly  relaying. 
 Pick the cells off the older pupal brood, look for similarity of  age.  If 
its patchy, either the queen is a spotty layer, or the colony is  losing 
brood during development.
 
Jerry

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