>> The bees clustered on the aged, drawn comb and were very sparse on the newly drawn comb.
I think I know what it is. You went from a tall nest where the queen laid in the new comb to a horizontal one. The coverage and thermal dynamics are quite different bet. the two. You need a lot more bees in the horizontal deep to cover the same amount of brood!
The bees build up more efficiently in vertical/narrow hives. The new comb is a much more secondary consideration. The best time to do this sort of re-configuration is when there is a lot of emerging brood.
Waldemar
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