PLB>My reading is that they are positing bee behavior as the main factor; mite behavior is not implicated in this work. 

Thanks- I understand that bee behavior is the main factor. My question was more about the section on "Mite reproduction in singly infested cells" where they suggest the following.

"Our results therefore suggest that the rate of Varroa mite reproductive failure has increased over time. Infertility rates are also higher than those reported for Varroa tolerant Cape honey- bees, on which mites remained infertile in 26% of both worker and drone cells (Allsopp, 2006). This could reflect the effect of selection on savannah honeybee colonies that has resulted in a reduction of mite population growth". 

Revising my question - how might selection on honey bees result in the reduction of mite fertility? 

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