> One would wonder, though, about colonies left packed in singles

The singles in question were one story Dadant hives, much bigger than Langstroth/Root hives

Dadant hive body: over-all measurements are 20 x 18 1⁄2 x 11 1⁄2 inches , with 11-12 frames 11 1/4 by 17 3/8 inches .

PLB

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