6 apiaries, Italian vs. Russian colonies, sun vs. shade, stocks  
separate or "comingled".  "Mite bomb" possibilites in some of the  
apiaries are described.  See Fig. 1 and 2.  Note that both Russian and  
Italian colonies did show increases in mite numbers even when the  
stocks were in separate apiaries.

https://www.ars.usda.gov/SP2UserFiles/Place/64133000/PDFFiles/401-500/417-Rinderer--The%20Effects%20of%20Co-mingled.pdf

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