I like to start with the simplest things first. In this situation i would have wondered if the mother had extroverted nipples that got pushed in with a tight or new bra causing a ring of moisture that never dried out. Simple irritation. ointments, etc. just continue to hold the moisture in especially if she didn't dry her nipples well before closing her bra. 
Vicky York, IBCLC, CPD
Oregon



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