Lindauer (1956a, b) showed that A. florea normally dance on a horizontal plain.
The experiment of rotating the comb so that the crown is upside down and therefore
no longer horizontal caused severely disturbed dancing. However, the bees will
automatically find a horizontal floor for dancing. For example, if the crown is
upside down, then the bees simply dance on the lower edge of the comb, which was
recently turned upward. If the crown is covered by other objects, the dance either
becomes disorientated or bees perform dances on the horizontal plane on top of
objects covering it. 

Like in a swarm cluster, the dwarf bees successfully dance in a vertical plane with a
similar rule: the dancers on the top of the cluster point directly to the goal, whereas
the dancers on the curtain are on the vertical plane and, instead of referring to
gravity like other cavity-nesting and giant bees do, the dance is orientated directly
towards the goal (Dyer and Seeley 1985a).

Source
Hepburn, H. R., & Radloff, S. E. (Eds.). (2011). Honeybees of Asia. Springer Science & Business Media.

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