Despite the fact that symmetry is common in nature, it is rarely perfect. Because
there is a wide range of phenotypes which differs from the average one, the asymmetry
should increase along with deviation. Therefore, the aim of this study was
to assess the level of asymmetry in normal individuals as well as in phenodeviants
categorized as minor or major based on abnormalities in forewing venation in honey
bees. Shape fluctuating asymmetry (FA) was lower in normal individuals and minor
phenodeviants compared with major phenodeviants ...

... The correlation between wing size and wing asymmetry
was negative, indicating that smaller individuals were more asymmetrical. The high
proportion of phenodeviants in drones compared with workers and queens confirmed
their large variability. 

The relationship between asymmetry, size and unusual venation in honeybees (Apis mellifera)
S. Łopuch and A. Tofilski
Department of Pomology and Apiculture, Agricultural University,
29 Listopada 54, 31-425 Krakow, Poland

[comment: phenodeviants? That's a new one for me. I consulted the Ngram Viewer and learned that the word first appeared in English in 1950 and was used a lot in the 1960s. It fell way off after 1970 and its use plummeted after 2000. Perhaps the translators found it in an old Polish to English dictionary]

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