https://pursuit.unimelb.edu.au/articles/feral-honeybees-key-to-controlling-deadly-parasite
Key take-away IMHO:
In our research, we found that feral honeybee colonies are distributed with remarkable uniformity.
We developed agent-based models of the mating system (computer models based on individual bees), to work out how the level of genetic diversity observed in the brood translated to densities of feral colonies in the area surveyed.
Using these models, our mid-range estimates indicate that there could be between 0.3 and 0.9 colonies per square kilometre in the vicinity of the Port of Newcastle.
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