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Date:
Sat, 29 Feb 2020 16:34:57 -0500
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Informed Discussion of Beekeeping Issues and Bee Biology <[log in to unmask]>
Subject:
Re: role of body size dimorphism in drones
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Informed Discussion of Beekeeping Issues and Bee Biology <[log in to unmask]>
From:
Frank Wright <[log in to unmask]>
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Here is a fairly recent study on the topic that appears to have decent controls.  

https://academic.oup.com/beheco/article/21/3/520/219340

This is the gist of their conclusions:  

Small males obtained approximately half the number of matings of normal-sized males, and those small males that did mate also obtained only 61% of the paternity of normal-sized males, which matches well with data showing that they have only 63% as many spermatozoa 

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