I should add a disclaimer: This is just my personal opinion, based on the
results of peer-review literature:
https://journals.plos.org/plosone/article?id=10.1371/journal.pone.0150362
*We hypothesized that the persistence of these wild colonies is due in part
to their habits of nesting in small cavities and swarming frequently.*
*...Colonies in small hives swarmed more often, had
lower Varroa infestation rates, had less disease, and had higher survival
compared to colonies in large hives. These results indicate that the
smaller nest cavities and more frequent swarming of wild colonies
contribute to their persistence without mite treatments. *
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