> Add to that  what actually triggers supercedure?  Read in many places its "lack of queen pheromone"  not exactly true because we know packages with fresh queens do it also. Are there multiple reasons? ... Interesting questions,   What could we do with the answers that would help us? 

I have been reading everything I can find on this topic for ten years, and have written numerous articles on the subject, published in the ABJ. The idea that "lack of queen pheromone" governs this things went out decades ago, it is far more interesting than that. But the sense I get is that the average beekeeper doesn't really care about the regulatory systems in the colony. He or she just wants more bees, more honey, more money. So, it's a conversation that doesn't really go anywhere.

> A trademark of eusocial insect species is reproductive division of labor, in which workers forego their own reproduction while the queen produces almost all offspring. The presence of the queen is key for maintaining social harmony, but the specific role of the queen in the evolution of eusociality remains unclear. A long-discussed scenario is that a queen either behaviorally or chemically sterilizes her workers. However, the demographic and ecological conditions that enable such manipulation are unknown.

Olejarz, J., Veller, C., & Nowak, M. A. (2017). The evolution of queen control over worker reproduction in the social Hymenoptera. arXiv preprint arXiv:1702.04873.

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