Obviously, bees living out in the open will benefit from hair trigger defensiveness, as also exhibited by Apis dorsata -- or the tendency to flee, as in Apis florea. I think each of these species exhibits behaviors that are inextricably linked with their nesting habits. Dorsata and the African bees are continually molested by marauders. 

  Seems to me,  the simple detail of primates such as Monkeys means that only the very meanest could survive in Africa, (most primates who like honey)??   No clue which came first,  but in terms of natural selection,  chimps and baboons would certainly push the issues.  Both are well know to like honey.

Charles

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