More from the aforementioned paper:
Variation in relative fitness is not sufficient to
ensure permanent adaptation; there must also be enough
variation in absolute fitness to encompass at least one
viable type within a population.
The most realistic prediction
might be that there will be many species, especially very
abundant species, that will successfully adapt to a gradually
deteriorating environment, but there will be many more that
will not. We have scarcely begun to investigate the evolutionary
dynamics of populations adapting to deteriorating
environments, however.
PLB
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