there is no essential difference between the
hive bees and the flying bees in terms of number of Varroa mites.
Only the water-canying bees were considerably less infested.
But in drones, the situation is different: a considerably greater
number of drones in the hive were found to be infested (41.9%) as
compared to the flying drones (5.2%) Of the drones caught at the drone
congregation area (6.1%) And consequently, the average amount of
infested drones (18.2%) is much greater than that of worker bees (3.6%)

SPREAD OF VARROA JACOBSON! FROM ONE COLONY TO ANOTHER. Apiacta 2, 1981


> The remarkable rate at which varroa has spread throughout most of the world is due both to the particular biology of the mite and to the nature of modern beekeeping. In eastern Europe and West Germany the rate of natural movement of the mites has been estimated at about 3 km per year. 

MITE PESTS OF HONEY BEES. Ann. Rev. EntomoL 1982. 27:229-52

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