Recent work concludes that deformed wing virus replicates in varroa  
mites and without varroa, the symptoms would not appear

 > The correlation between DWV replication in mites and the  
development of wing deformity (and conversely, the correlation between  
absence of replication in mites and absence of clinical symptoms in  
bees) suggested that, although DWV transmission by V. destructor is a  
prerequisite for the development of crippled wings, a crucial factor  
is the replication of DWV in the mite prior to transmission.  We  
conclude that the development of crippled wings not only depends on  
DWV transmission by V. destructor but also on viral replication in V.  
destructor and on the DWV titre in the parasitizing mites.

Deformed wing virus: replication and viral load in mites (Varroa  
destructor)
Sebastian Gisder, et al
Journal of General Virology (2009), 90, 463–467

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