Pollen stored in comb cells of the hive has been reported as a bee
pathogen reservoir. Viable spores of Ascosphaera apis (chalk brood)
have been shown to survive in ''bee bread'' for at least 12 months.
Gilliam et al. have shown that the pollen stored in comb cells of the
hive differs biochemically from the corbicular pollen. Nosema spores'
viability for months in bee bread could be an important
epidemiological factor in the transmission of pathogens to the next
bee generation. Nosema spores in pollen can be a source of nosemosis
for the colony.

-- 
"Detection of infective Nosema ceranae (Microsporidia) spores in
corbicular pollen of forager honeybees"
Mariano Higes, et al

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