We found that we observed small bees in spring before we began feeding 
upplement, but that they became much less common after we made it standard 
ractice.

Could it be that the small bees in spring were raised in small cells in the core of the brood nest, but as the brood nest expanded they moved onto areas with bigger cells?  I think Dace Cushman's website may still have some information relating to measurements fo plaster casts I took of cells almost a decade ago.

Chris

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